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An association with a community-wide standard parameter has not yet been made.
code describing colony activity over previous year: D = death; I = fission (one colony splits into two or more colonies); M = fission and fusion (one colony undergoes both fission and fusion in one year); N = new (code given to the measurement of a colony in the first year of the project); R = recruit (new; small colony); S = survival (colony persists from year t to t+1); U = fusion (two physiologically distinct colonies merge into a single colony and take the name of the larger colony); X = phoenix (colony was present in year t-1; not visible in year t; but reappeared in year t+1)